The option that contains all of the other options listed is chromosome.
Here's the reasoning:
- A gene is a segment of DNA that is located on a chromosome.
- A chromatid is one half of a duplicated chromosome, so each chromosome consists of two chromatids after DNA replication.
- A centromere is the region of a chromosome that joins two sister chromatids.
Since a chromosome includes genes, chromatids (when duplicated), and a centromere, it encompasses all the other listed options.