For the excerpt you've provided, I would use an informative and slightly nostalgic tone.
Reasoning for this Tone:
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Informative: The content presents factual historical information about the unique use of glowworms during WWI. An informative tone would help convey the significance and necessity of this adaptation in a clear and engaging way, ensuring that the audience understands the practical application of glowworms in a challenging context.
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Nostalgic: The mention of glowworms in the context of war evokes a sense of creativity and resourcefulness from that era. A hint of nostalgia in the tone could capture the ingenuity of soldiers who found unconventional solutions in difficult circumstances, allowing listeners to appreciate the historical significance and human spirit behind such actions.
By combining these tones, the reading can engage the audience while effectively delivering the historical context and the intriguing use of glowworms.