The statement "Earth's many species of fungi act as decomposers of matter" is also a strong candidate for identifying a central idea of the paragraph, as it highlights an important role that fungi play in ecosystems. However, the paragraph emphasizes the interconnected underground networks of mycelia and their relationship with tree roots more prominently.
While both statements reflect significant points made in the paragraph, "Fungi exist as a web of threads connected underground" closely aligns with the primary focus on those networks and their ecological importance.
Thus, both statements are valid, but if you need to choose the one that captures the overarching theme presented in the paragraph, "Fungi exist as a web of threads connected underground" would still be the better choice.