In the combining terms you provided, the roots can be underlined as follows:
- splen/o (where "splen" is the root meaning "spleen")
- hyster/o (where "hyster" is the root meaning "uterus")
The roots are typically the main part of the word that conveys its primary meaning, while the combining form includes a vowel (in this case, "o") to facilitate pronunciation when combined with other word parts.