The best explanation for the difference in pigmentation observed in Himalayan rabbits, where one rabbit has pigmentation on its extremities and the other has none, is b) Environment affects the expression of the genes for that trait.
In Himalayan rabbits, the gene responsible for fur pigmentation is temperature-sensitive. The gene is expressed differently depending on the temperature; cooler areas of the rabbit's body (like the ears, nose, and feet) allow for pigment production, while warmer areas (like the body) do not. This is a classic example of how environmental factors can influence gene expression.