The main selective pressure in this scenario is D) Fish population.
The decline in the fish population directly affects the long-legged birds, which rely on fish as their primary food source. As the availability of fish decreases, the long-legged birds would face increased competition for food, leading to greater selection pressure on them. In contrast, the short-legged birds that feed on insects would not be affected by the decline in fish, as their food source remains stable. Thus, the decline in the fish population creates a significant pressure that impacts the long-legged birds directly.