John Allen’s perspective on the evolution of cells with chloroplasts and mitochondria is closely related to the endosymbiotic theory. According to this theory, complex cells (eukaryotes) arose from a symbiotic relationship between early simple cells (prokaryotes) and certain organelles that became incorporated into them.
Based on your description, it seems that Allen believes that both chloroplasts and mitochondria played a significant role in the evolution of more complex cells through beneficial interactions. The best option that reflects this idea is:
a) mitochondria engaged in reactions that benefited simple cells and recent complex cells.
This suggests that mitochondria provided benefits to both early simple cells and have continued to provide advantages to more complex cells throughout evolution, aligning with the endosymbiotic theory.