The most likely keystone species in the options provided is wolves in Yellowstone National Park, which eat elk, who graze on willows, on which beavers depend to build their dams.
Keystone species are those that have a disproportionately large effect on their environment relative to their abundance. Wolves play a critical role in controlling the population of elk, which in turn affects the vegetation in the ecosystem. This cascading effect supports other species in the ecosystem, making wolves a classic example of a keystone species.