The most likely keystone species among the options provided is wolves in Yellowstone National Park, which eat elk, who graze on willows, on which beavers depend to build their dams.
Keystone species have a disproportionately large impact on their environment relative to their abundance. Wolves in this scenario play a critical role in controlling the elk population, which in turn affects the vegetation, specifically willows. This cascading effect demonstrates how wolves help maintain the ecological balance within their ecosystem, making them a classic example of a keystone species.