The most likely statement is: The proportion of short-tailed squirrels may have decreased due to decreased food availability.
This is because as long-tailed squirrels are better adapted for jumping and balancing in trees, a decrease in food availability (for example, if food is more accessible in trees) could lead to a decline in the proportion of short-tailed squirrels, who may not be as efficient in navigating their environment as their long-tailed counterparts. This would result in a relative increase in the long-tailed population over time.