Based on the behaviors and causes related to swim bladder issues in koi, here are the matches:
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Stays on the surface and can't swim well or hide: The koi has a parasitic infection, like a worm, that prevents air from entering the swim bladder.
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Sinks to the bottom and can't fill the swim bladder or feed: The koi is stressed by dirty water with too much waste and too many nitrates, the swim bladder can't empty correctly.
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The koi eats food or insects that are too large, blocking the air intake: The koi experiences difficulty in filling the swim bladder.
These associations help in identifying the underlying environmental issues or physical ailments affecting koi behavior and their swim bladder functionality.