Among the four main types of biomolecules—carbohydrates, lipids, proteins, and nucleic acids—lipids hold the most energy.
Specifically, lipids such as fats and oils are highly energy-dense due to their long hydrocarbon chains. They provide about 9 calories of energy per gram, which is more than double the energy provided by carbohydrates and proteins, both of which provide about 4 calories per gram. This high energy content makes lipids an important energy source for organisms.