The amplitude of the depolarization at the sensory receptor (R1) is primarily determined by the strength of the stimulus applied to the sensory receptor. In sensory neurons, a stronger stimulus typically generates a larger receptor potential, leading to a greater depolarization. This depolarization can then influence the frequency of action potentials generated, but the initial amplitude of depolarization is directly related to the stimulus strength.
The other options mentioned relate to aspects of neuronal signaling but do not directly determine the initial amplitude of the depolarization at the sensory receptor itself.